Consider a lake with some fish attractive to fishermen. We wish to model the fish-fishermen interaction. Fish Assumptions: i. Fish grow logistically in the absence of fishing. ii. The presence of fishermen depresses fish growth at a rate jointly proportional to the fish and fishermen populations. Fishermen Assumptions: 1\. Fishermen are attracted to the lake at a rate directly proportional to the amount of fish in the lake. i. Fishermen are discouraged from the lake at a rate directly proportional to the number of fishermen already there. (a) Formulate, analyze, and interpret a mathematical model for this situation. (b) Suppose the department of fish and game decides to stock the lake with fish at a constant rate. Formulate, analyze, and interpret a mathematical model for the situation with stocking included. What effect does stocking have on the fishery?

Short Answer

Expert verified
Stocking the lake increases the fish population by a constant rate, potentially leading to a higher equilibrium population.

Step by step solution

01

Define Variables

Let time be denoted by t, fish population by F(t), and fishermen population by M(t). These variables will help in constructing the equations.
02

Fish Population Dynamics

Use the logistic growth model for fish without fishing: \ \( \frac{dF}{dt} = rF \left(1 - \frac{F}{K}\right) \) where r is the intrinsic growth rate and K is the carrying capacity. Incorporate the negative impact of fishermen: \ \( \frac{dF}{dt} = rF \left(1 - \frac{F}{K}\right) - aFM \) where a is the rate at which fishermen depress fish growth.
03

Fishermen Population Dynamics

Fishermen are attracted to the lake by fish at a rate proportional to the fish population: \ \( bF \). They are discouraged at a rate proportional to their own population: \ \( dM \). Combining both effects: \ \( \frac{dM}{dt} = bF - dM \).
04

Mathematical Model Formulation

Combine the equations for both fish and fishermen populations: \ \( \frac{dF}{dt} = rF \left(1 - \frac{F}{K}\right) - aFM \ \frac{dM}{dt} = bF - dM \)
05

Impact of Stocking on Fish Population

To model the constant stocking rate, denote the stocking rate as S. Modify the fish population equation to include stocking: \ \( \frac{dF}{dt} = rF \left(1 - \frac{F}{K}\right) - aFM + S \). The fishermen population equation remains the same.
06

Stocked Model Formulation

The mathematical model with constant stocking is: \ \( \frac{dF}{dt} = rF \left(1 - \frac{F}{K}\right) - aFM + S \ \frac{dM}{dt} = bF - dM \)
07

Interpretation of Stocking Effect

Stocking increases the fish population by a constant rate S, which can help counteract the negative impact of fishing, potentially leading to a higher equilibrium fish population.

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Key Concepts

These are the key concepts you need to understand to accurately answer the question.

logistic growth
In the study of population dynamics, logistic growth is a fundamental concept. It describes how a population grows rapidly when it is small, slows as it approaches the environment's carrying capacity, and eventually stabilizes. The equation that represents logistic growth is:
\[ \frac{dF}{dt} = rF \bigg(1 - \frac{F}{K}\bigg) \] where:
  • \( F \) represents the fish population,

  • \( r \) is the intrinsic growth rate,

  • \( K \) is the carrying capacity, the maximum population size that the environment can sustain.

This model shows that when the population size \( F \) is far below the carrying capacity \( K \), the growth rate is high. As \( F \) approaches \( K \), the term \( 1 - \frac{F}{K} \) becomes smaller, slowing the growth rate until growth stops altogether when the population size hits the carrying capacity.
population dynamics
Population dynamics explores how and why the numbers of individuals in populations change over time. In this fish-fishermen interaction model, dynamics are influenced by multiple factors:
  • Fish population grows logistically without fishing.

  • The presence of fishermen negatively impacts fish growth at a rate proportional to both fish and fishermen populations.

  • Fishermen are attracted to and discouraged from the lake based on the fish population and the number of other fishermen, respectively.

Combining these, the fish and fishermen populations are described by the following system of differential equations:
\[ \frac{dF}{dt} = rF \bigg(1 - \frac{F}{K}\bigg) - aFM \] \[ \frac{dM}{dt} = bF - dM \] where:
  • \( r \) is the fish intrinsic growth rate,

  • \( K \) is the carrying capacity,

  • \( a \) is the rate at which fishermen depress fish growth,

  • \( b \) is the rate at which fishermen are attracted to the lake by fish,

  • \( d \) is the rate at which fishermen leave the lake.

This model highlights the dynamic and interdependent nature of the fish and fishermen populations.
constant rate stocking
Introducing a constant rate of stocking fish into the lake alters the dynamics. Stocking adds a fixed number of fish to the population over time, helping to mitigate the negative effects of fishing.
The modified fish population equation with constant rate stocking \( S \) is:
\[ \frac{dF}{dt} = rF \bigg(1 - \frac{F}{K}\bigg) - aFM + S \] Here, \( S \) represents the number of fish added to the lake per unit time.

The inclusion of stocking can increase the equilibrium fish population compared to when there's no stocking. This means that even with fishing pressures, the fish population can be maintained at a higher level due to the continuous addition of new fish.
The fishermen population remains influenced only by the fish population and their own numbers, hence the equation
\[ \frac{dM}{dt} = bF - dM \] remains unchanged. Stocking compensates for the fish removed by fishermen, which can result in a stabilized fish population that supports ongoing fishing activities.

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Most popular questions from this chapter

Species may derive mutual benefit from their association; this type of interaction is known as mutualism. May (1976) suggests the following set of equations to describe a possible pair of mutualists: $$ \frac{d N_{1}}{d t}=r N_{1} \frac{1-N_{1}}{\kappa_{1}+\alpha N_{2}}, \quad \frac{d N_{2}}{d t}=r N_{2} \frac{1-N_{2}}{\kappa_{2}+\beta N_{1}} $$ where \(N_{i}\) is the population of the ith species, and \(\alpha \beta<1\). (a) Explain why the equations describe a mutualistic interaction. (b) Determine the qualitative behavior of this model by phase-plane and linearization methods. (c) Why is it necessary to assume that \(\alpha \beta<1 ?\)

Anderson and May (1979) describe a model in which the natural birth and death rates, \(a\) and \(b\), are not necessarily equal so that the disease-free population may grow exponentially: $$ \frac{d N}{d t}=(a-b) N $$ The disease increases mortality of infected individuals (additional rate of death by infection \(=\alpha\) ), as shown in the accompanying figure. In their terminology the population consists of the following: $$ \begin{aligned} &X=\text { susceptible class }(=S) \\ &Y=\text { infectious class }(=I) \\ &Z=\text { temporarily immune class }(=R) \end{aligned} $$ (a) Write equations describing the disease. (b) Show that the steady-state solution representing the presence of disease is given by Applications of Continuous Models to Population Dynamics 265 $$ \bar{X}_{2}=\frac{\alpha+b+v}{\beta}, \quad \bar{Y}_{2}=\frac{r}{\alpha} N, \quad \bar{z}_{2}=\frac{r}{\alpha}\left(\frac{\nu}{b+\gamma}\right) N_{2} $$ where $$ N_{2}=\frac{\alpha(\alpha+b+\nu)}{\beta(\alpha-r[1+\nu /(b+\gamma)])}, \quad \text { and } \quad r=a-b $$ (c) The steady state in part (b) makes biological sense whenever $$ \alpha>r\left(1+\frac{\nu}{b+\gamma}\right) $$ Interpret this inequality. (Note: Anderson and May conclude that disease can be a regulating influence on the population size.)

Show that in an SIR model with disease fatality at rate \(\eta\) the disease will always eventually disappear.

Beddington and May (1982) have proposed the following model to study the interactions between balcen whales and their main food source, krill (a small shrimp-like animal), in the southem ocean: $$ \begin{gathered} \dot{x}=r x\left(1-\frac{x}{K}\right)-a x y \\ \dot{y}=s y\left(1-\frac{y}{b x}\right) \end{gathered} $$ Here the whale carrying capacity is not constant but is a function of the krill population: $$ K_{\text {whales }}=b x $$ Analyze this model by determining the steady states and their stability; include a phase-plane diagram.

(a) Suppose a one-time fishing expedition reduced the prey population by \(10 \%\) of its current level. What does the Lotka-Volterra model predict about the subsequent behavior of the system? (Note: this prediction is one of the most objectionable features of the model and will be dealt with in a. later chapter.) (b) Now consider the situation in which there is a constant level of fishing in which both prey and predatory fish are caught and removed at rates proportional to their densities, \(\phi x\) and \(\phi y\). Compare this to the situation in the absence of fishing, and show what Volterra concluded about d'Ancona's observation. (For one treatment of this problem see Braun, 1979 ; a more advanced mathematical treatment can be found in Brauer and Soudack, 1979.)

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